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thanwasthatofNorthAmerica比北美的什么热?为什么要加WAS?ManymoreofthespeciesoflargermammalsthanofthespeciesofsmallermammalslivinginNorthAmericainthatperiodhadoriginatedinclimatesthatwerewarmerthanwasthat

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than was that of North America 比北美的什么热?为什么要加WAS?
Many more of the species of larger mammals than of the species of smaller mammals living in North America in that period had originated in climates that were warmer than was that of North America before the end of the Pleistocene period.

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than这个比较句式,肯定是同类事物的比较,所以这里的that指代climatewas也是同样道理,比较的两部分形式做到一致,前面有“were”,所以后半句也要加was相当于: had originated in climates that were warmer than was ...